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How do Principal Investigators register on the NIH eRA Commons website?

It can be done easily via University of Maryland (ORAA).

All PIs must be registered on NIH's eRA Commons System before a proposal can be submitted to NIH. This registration is done by the University of Maryland (by ORAA), not through NIH. To register, go to ORAA's website and provide your UMD username and password:

http://www.ora.umd.edu/esubmissions/nih